Lee Kong Chian Natural History Museum

Benito C. Tan


Dr. Ben C. Tan studies the taxonomy, systematics, evolution and conservation of East Aisatic and Malesian moss diversity for more than 30 years. He also is interested in the taxonomy of Philippine fern flora, especially the group of Selaginella. He has conducted numerous expeditions to collect and document the diversity of mosses in many countries in Asia and North America. As a result thereof, he has published more than 200 technical publications in various international journals and written several chapters in books on Asiatic mosses.  
Dr. Tan is a member of the Committee on Nomenclatures on Bryophyta of the International Association of Plant Taxonomists (IAPT), and a member of the Committee on Conservation of Mosses of International Association of Bryoplogists (IAB). He is an adjunct researcher at RMBR of NUS and at the University Herbarium of the University of California at Berkeley. Dr. Tan is member of Editorial Board of several international bryological journals.

He is currently an Adjunt Professor at the Department of Biological Sciences at the National University of Singapore.


Tan, B.C. and  Ho B.-C. 2010. Moss power, pp. 26–35. In: D. Yeo C. J., L. K. Wang and K. Lim K. P. (eds). Private Lives, an Exposé of Singapore Freshwaters.  RMBR-NUS and Exxon Mobil.

Li, F.-W., B. C. Tan, V. Buchbender, R. C. Moran, G. Rouhan, C.-N. Wang and D. Quandt. 2009. Identifying a mysterious aquatic fern gametophyte. Plant Systematics and Evolution 281:1-4, 77-86 (Online publication date: 1-Sep-2009).

Wang, Y.-M.  L. J. Harrison and B. C. Tan. 2009. Terpenoids from the liverwort Chandonanthus hirtellus. Tetrahedron 65: 4035–4043.

Wongkuna K., K. Santanachote and B. C. Tan. 2009. Miscellaneous observation on Fissidens in Thailand with five new species records. Cryptogamie, Bryologie 30: 301-309.

Hedenaes, L., B.C. Tan and Y. Chang. 2008 (2009). Morphological and molecular data suggest different evolutionary relationships among species of Acroporium and closely related genera (Sematophyllaceae), pp. 259-276. In: Mohammed, H., Baki, B.B., Nasrulhaq-Boyce, A. and Lee, P.K.Y.(eds.). Bryology in the New Millenium. University of Malaya, Kuala Lumpur.  

Linis, V.C. and B.C. Tan. 2008 (2009). Progress of studies on phytogeography and biodiversity of the Philippine moss flora from 1991to 2006, pp. 13-22. In: Mohammed, H., Baki, B.B., Nasrulhaq-Boyce, A. and Lee, P.K.Y.(eds.). Bryology in the New Millenium. University of Malaya, Kuala Lumpur.  

Tan, B.C.and Boon-Chuan Ho. 2008. A Guide to the Mosses of Singapore. Science Centre, 149 pp.

Tan, B.C. and B.-C. Ho. 2008. Bryophytes. In: G.W.H. Davison, P.K.L. Ng & H.C. Ho (eds.), The Singapore Red Data Book:Threatened Plants & Animals of Singapore, 2nd ed., pp. 17–19, 209. The Nature Society (Singapore).

Tan, B.C. 2007. Are there indicator bryophyte species for lowland rainforests in Asian tropics?  pp. 109-112. In: R. Stuebing, J. Unggang. J. Ferner, et al., Proceedings of the Regional Conference of Biodiversity Conservation in Tropical Planted Forests in Southeast Asia, 15-18 January 2007.  Sarawak Forestry Corporation and Grand Perfect Sdn Bhd, Kuching. 

Tan, B.C., S. Piippo and D.H. Norris. 2007. Bryophytes of Papua New Guinea: Their Diversity, Ecology and Conservation, pp. 320-334. In: A.J. Marshall and B.M. Beehler (eds.). The Ecology of Papua, Part One. The Ecology of Indonesia Series, Vol. VI. Periplus. 

Tan, B.C., B.-C. Ho, V. Linis, E.A.P. Iskandar, I. Nurhasanah, L. Damayanti, Sri Mulyati and I. Haerida. 2006. Mosses of Gunung Halimun National Park, West Java, Indonesia.  Reinwardtia 12: 205-214. 

Cao, T., R. Zhu, B.C. Tan, S. Guo, C. Gao, P. Wu and X. Li. 2006.  A report of the first national red list of Chinese endangered bryophytes. Journal of Hattori Botanical Laboratory 99: 275-295.

Chang Ying, Chan W.K, S.W. Graham, and B.C. Tan. 2006.  Molecular evidence for the systematic positions of two enigmatic mosses: Pterogonidium pulchellum (Sematophyllaceae, Musci) and Piloecium pseudorufescens (Myuriaceae, Musci). Canadian Journal of Botany 84 (3): 501-507.

Guo, S.-L., B.C. Tan and V. Virtanen. 2006. Taxonomic and morphometric comments on Macromitrium blumei, M. zollingeri and M. annamense (Orthotrichaceae, Bryophyta).  Nova Hedwigia 82: 467-482.

Ho, B.C., B.C. Tan and N.S. Hernawati. 2006. A checklist of mosses of Sumatra, Indonesia. Journal of Hattori Botanical Laboratory 100: 143-190.

Leonardia, A.A.P., P. Kumar and B.C. Tan. 2006. Development of microsatellite markers for the tropical moss, Acanthorrhynchium papillatum. Molecular Ecology Notes 6: 396-398.   

Lim, T.B., R. Xu, B.C. Tan and J.P. Obbard. 2006. Persistent organic pollutants in moss as bioindicators of atmosphere pollution in Singapore. Chemosphere 64: 596-602.

O’Shea, B.J. and B.C. Tan. 2006. British Bryological Society Expedition to Mulanje Mountain, Malawi. 14. Sematophyllaceae (Bryopsida): Part 1. Journal of Bryology 28: 360-362.  

Suleiman, M., H. Akiyama and B.C. Tan. 2006. A revised catalogue of mosses reported from Borneo. Journal of Hattori Botanical Laboratory 99: 107-183.

Tan, B.C. 2005. New species records of Sri Lankan mosses. Raffles Bulletin of Zoology, Supplement 12: 335-338.

Tan, B.C. and G. Dauphin. 2005. A new genus and species, Pilosimitra buckii (Sematophyllaceae, Musci) from Costa Rica. Tropical Bryology 26: 19-22.

Gradstein, S.R., B.C. Tan, R.-L. Zhu, B.-C. Ho, C. King S.-H., C. Drűbert and R. Pitopang. 2005. A catalogue of the bryophytes of Sulawesi, Indonesia. Journal of Hattori Botanical Laboratory 98: 213-257.  

Li, Z.-J. and B.C. Tan. 2005. A review of the species diversity of Selaginella in Fujian Province of China. Acta Phytotaxonomica Sinica 43: 50-59.  

Tan, B.C. and Chang Ying. 2004. Molecular contribution to the systematic position of Mastopoma scabrifolium (Broth. in Moell.) B.C. Tan & Tran Ninh (Sematophyllaceae, Bryopsida). Cryptogamie, Bryologie 25: 301-308.    

Tan, B.C., B.-C. Ho and B. K.-B. Shea. 2004. Two new moss species, Trichosteleum fleischeri and Splachnobryum temasekensis, from Singapore.  Journal of Hattori Botanical Laboratory 96: 223-230.

Tan, B.C. and Tan Swee-Hee. 2004. Country Report: Singapore – Its biodiversity, conservation and resource management, pp. 68-75.  In: Proceedings of Chinese-ASEAN Workshop on Conservation and Biotechnology Application of Tropical Resources, held in October of 2004 at Haikou, China.

Hedenäs, L., B.C. Tan, I. Bisang and C. Hammarberg. 2004. Habitat preference in Acroporium (Sematophyllaceae) and related taxa in Peninsular Malaysia. Bryologist 107: 550-565.

Ignatov, M. T. Tsegmed, B.C. Tan, X.-L. Bai and V. Zolotov. 2004. Mosses of Gobi in Mongolia. Journal of Hattori Botanical Laboratory 96: 183-210.  

Ramsay, H.P., W.B. Schofield and B.C. Tan. 2004. The Family Sematophyllaceae (Bryopsida) in Australia. Part 2. Acroporium, Clastobryum, Macrohymenium, Meiotheciella, Meiothecium, Papillidiopsis, Radulina, Rhaphidorrhynchium, Trichosteleum and Warburgiella. Journal of Hattori Botanical Laboratory 95: 1-69.

Tan, B.C. 2003. Bryophytes (mosses). In: W.P. de Winter & V. B.Amoroso (eds.). Cryptogams: Ferns and fern Allies. Plant Resources of South East Asia (PROSEA)15(2): 193-200.

Tan, B.C. and Tran Ninh. 2003. Vu Quang and other Vietnam mosses collected by Tran Ninh, B.C. Tan and T. Pócs in 2002. Acta Acad. Paed. Agriensis, sectio Biologie, n.s. 24: 85-101.

Tan, B.C. 2002. The affinity of moss flora of Japan, Taiwan and the Philippines revisited: old problems, new insight and more questions. Acta Phytotax. Geobot. 53: 77-84.

Tan, B.C. and W. R. Buck. 2002. Pterogonidium pulchellum, a new moss record for Asia from Singapore. Folia Malaysiana 3: 166-170.

Farida binti Yusuf, B.C. Tan and I.M. Turner. 2003. What is the minimum area needed to estimate the biodiversity of pteridophytes in natural and man-made lowland forests in Malaysia and Singapore? Fern Gazette 17: 1-9.

Farida binti Yusuf, B.C. Tan and I. Turner. 2003. Species richness of pteridophytes in natural versus man-made lowland forests in Malaysia and Singapore, pp. 283-298. In: S. Chandra and M. Srivatava (eds.). Pteridology in the New Millennium.  Kluwer Academic Publishers, Netherlands.   

Lin, B.-J. and B.C. Tan. 2002 (2003). Hookeriaceae, pp. 3-34. In: Wu P.-C., M.R. Crosby nd S. He (eds.). Moss Flora of China, English version, vol. 6. Hookeriaceae–Thuidiaceae. Science Press and Missouri Botanical Garden (St. Louis).

Ramsay, H.P., W.B. Schofield and B.C. Tan. 2002. The family Sematophyllaceae (Bryopsida) in Australia. Part 1. Introduction, family data, key to genera and the genera Wijkia, Acanthorrhynchium, Trismegistia and SematophyllumJournal of Hattori Botanical Laboratory 92: 1-50.

Tan, B.C. and Choy Meng-Shyan. 2002. The taxonomy, phytogeography and conservation of the Fissidens flora of Singapore, with one new species, F. pseudoceylonensis. Journal of Bryology 24: 45-55.

Piippo, S. X.-L. He, A. Juslén, B.C. Tan, D.H. Murphy and T. Pócs.  2002. The hepatic and hornwort flora of Singapore.  Ann. Bot. Fenn.39: 101-127.  

Iwatsuki, Z. and B.C. Tan. 2001. Entodon challengeri (Par.) Card., the correct name for E. compressus sensu Mizushima and Crum and Anderson. Taxon 50: 1125-1128.  

Tan, B.C. 2000. The first moss exsiccata of Brunei Darussalam issued by the Raffles Mueum for Biodiversity Research, National University of Singapore. Raffles Bull. Zool. 48: 338.  

Tan, B.C. and T. Pocs. 2000. Biogeography and conservation of bryophytes, pp. 403-448. In: A.J. Shaw and B. Goffinet (eds.). Bryophyte Biology. Cambridge University Press.

Ignatov, M.S., B.C. Tan, Z. Iwatsuki and E. A. Ignatova. 2000. Moss flora of the Upper Bureya River, Russian Far East. J Hattori Bot. Lab. 88: 147-178. 

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